PARASITISM
Parasitism may be regarded as a special form of predation in which the predator
is much smaller than the prey and remains closely associated with it. Parasitism
is harmful to the prey organism and beneficial to the parasite. The concept
of parasitism seems obvious, but individual instances are often surprisingly,
difficult to distinguish from predation and from various other kinds of symbiosis.
Many instances of parasitism are well known; for example, vertebrates are
parasitized by members of many different phyla of animals and protists. Invertebrates
also have many kinds of parasites that live within their bodies. However,
bacteria and viruses are often not considered parasites, even though they
fit our definition precisely. Lice, which live on the bodies of vertebrates-mainly
birds and mammals-are normally considered parasites, but mosquitoes are not,
even though they draw food from the same birds and mammals in a similar manner,
because their interaction with their host lasts for only a short time. However,
mosquitoes are closely associated ecologi-callywith the animals from which
they draw blood. Mosquitoes also synchronize their diurnal and seasonal activities
closely with those of their hosts, so that the interrelationship is very close.
Many fungi and some flowering plants, too, are parasitic on other plants,
and a few are serious pests of crops.
Internal parasitism is generally marked by much more extreme specialization
than external parasitism, as shown by the many protist and invertebrate parasites
that infect humans. The more closely the life of the parasite is linked with
that of its host, (he more its morphology and behavior are likely to have
been modified during the course of its evolution. The same, of course, is
true of symbiotic relationships of all sorts. Conditions within the body of
another organism are different from those encountered outside and are apt
to be much more constant in every way. Consequently, the structure of the
parasite is often simplified, and unnecessary armaments and structures are
lost as it evolves.
The ecosystem is the most complex level of biological organization. Communities
are composed of the organisms present at a particular place, whereas ecosystems
include the nonliving factors interacting with these organisms. In ecosystems
there is a regulated transfer of energy and a controlled cycling of nutrients.
The individual organisms and populations of organisms in an ecosystem act
as parts of an integrated whole, adjust over time to their role in the ecosystem,
and relate to one another in complex ways that we only partly understand.
Despite their differences, all ecosystems regulate the flow of energy-ultimately
derived from the sun-and the cycling of nutrients. The earth is a closed system
with respect to the chemicals, but an open one in terms of energy. Collectively,
the organisms that occur in ecosystems regulate [he capture and expenditure
of that energy and the cycling of those chemicals. All organisms, including
humans, depend on the ability of a few other organisms-plants, algae, and
some bacteria in the case of carbon, and certain bacteria in the case of nitrogen,
for example-for the basic components of life.
As distinct functional units, different kinds of ecosystems have more or less
clearly recognizable boundaries, but they also intergrade into one another,
sometimes almost imperceptibly; boundaries then become arbitrary. Ecosystems
also change over time and slowly become modified into new ecosystems, whose
characteristics come to differ increasingly from those that preceded them.
Thus the com-ilex ecosystems of the tropical rain forests have changed gradually
in adapting to the articular conditions of temperature, seasonably, and soil
that typify these places. Fhe ecosystems of the tundra have developed in a
similar way, but in relation to the Unerent environmental conditions of the
far north. As the climate changes in a given ilace, the ecosystems that are
present there change along with it, as do the individual populations within
these ecosystems. By this process, the overall characteristics of the populations
gradually adjust to the new conditions. Not all ecosystems are natural; we
nay also speak of an ecosystem in an aquarium or in a cultivated field.


